WITH few exceptions, tooth shape among crocodyliform reptiles (Crocody
lia of traditional use) is rather uniform(1). We report here on the pr
esence of multicuspid molariform teeth in a remarkable new crocodylifo
rm from the Lower Cretaceous of China, which may represent the first k
nown herbivorous member of that group. The overall structure of these
teeth is very similar to that of the postcanine teeth of tritylodontid
synapsids and represents a particularly striking example of convergen
t evolution, It indicates back-to-front (proal) motion of the mandible
produced by the posterior pterygoid muscle during jam closing, much a
s in the extant tuatara, Sphenodon(2,3). Certain derived features indi
cate that the new Chinese crocodyliform is closely related to the Noto
suchidae from the Cretaceous of Gondwana(4). Its discovery thus casts
further doubts on claims(5) concerning an endemic Gondwanan tetrapod f
auna during the Cretaceous.