WITH nearly 4,000 living species(1), frogs are numerically the most su
ccessful of modern amphibian groups, Their distinctive anatomy, which
exhibits numerous unique features in both the axial and appendicular s
keletons(2-6), represents a major departure from the body plan of Pala
eozoic amphibians, We report here the discovery of the earliest known
frog, associated with caecilians that retained limbs(7), that exhibits
primitive characters but shares with later anurans such features as f
usion of the caudal vertebrae (urostyle), a rod-like ilium, and elonga
te hind limbs. The evolution of saltation in anurans entailed transloc
ation of the ilium below the sacral rib, recruitment of the primitive
tail musculature to transmit force from the hind limb to the axial ske
leton during a jump, and development of iliosacral mobility, We reinte
rpret the caudopelvic transition from Palaeozoic amphibians to modern
frogs with reference to Triadobatrachus massinoti, an Early Triassic a
mphibian that possesses some frog-like features(8). The Early Jurassic
age and Laurasian provenance of the new frog support the hypothesis(4
) that the widespread occurrence of primitive extant anurans reflects
an ancient Pangaean distribution.