The distribution of charophyte assemblages in the Purbeck and Wealden
sequence of southern England has been established from borehole sample
s from the Weald and from outcrop material collected in Dorset, Wiltsh
ire and on the Isle of Wight. Of the twenty-one taxa represented, thre
e are new: Globator rectispirale, Clypeator britannicus and Sphaerocha
ra andersonii; three new combinations are proposed: Globator praecurso
r, Globator protoincrassatus and Atopochara triquetra. The Chinese Val
anginian species Flabellochara xiangyunensis is recognized for the fir
st time in Europe. In the context of the phylogeny of the Family Clava
toraceae, G. rectispirale represents the Jurassic ancestor of the Glob
ator lineage and a separate origin is suggested for both Flabellochara
and Clypeator. The correlations established with the Tethyan realm lo
cate the Jurrassic-Cretacous boundary within the Lulworth Formation of
the Purbeck Limestone Group; in this context, the whole 'Purbeck' seq
uence of Swindon (Wiltshire) is attributed to the Upper Tithonian. The
distribution of Clavatoraceae indirectly confirms the contemporaneity
of the Boreal Galbanites kerberus and Titanites anguiformis with the
Tethyan 'Durangites' ammonite zones. For the Wealden Supergroup, the c
harophyte data affirm the Hauterivian-Barremian boundary at the base o
f the upper division of the Weald Clay and the Upper Barremian is iden
tified at the base of the Vectis Formation of the Isle of Wight.