SYNOPSIS OF THE TRIBE BOCAGEEAE (ANNONACEAE), WITH REVISIONS OF CARDIOPETALUM, FROESIODENDRON, TRIGYNAEA, BOCAGEA, AND HORNSCHUCHIA

The tribal name Bocageeae Endlicher is reestablished and the tribe is circumscribed on the basis of solitary internodal ebracteate pedicels that are articulated at the base, and pollen shed in polyads of eight or more grains. Septate anther locules, large pollen size, and seed ap pendages are prevalent in the tribe. Intectate pollen with free-standi ng columellae, rare in Annonaceae, occurs not only in the genus Trigyn aea but also in some species of Bocagea and Hornschuchia. As defined h ere, the Bocageeae include seven neotropical genera: Cymbopetalum, Por celia, Bocagea, Cardiopetalum, Froesiodendron, Hornschuchia, and Trigy naea. The latter five genera are revised and the treatments include te n new species: Cardiopetalum plicatum, Froesiodendron urceocalyx, Horn schuchia lianarum, H. santosii, H. leptandra, Tripynaea cinnamomea, T. lanceipetala, T. triplinervis, T. lagaropoda, and T. axilliflora, all from tropical South America. A new combination, Froesiodendron longic uspe, changes the rank of that taxon from subspecies to species. Cardi opetalum surinamense is removed from Froesiodendron on and reassigned to Cardiopetalum on the basis of its connate petals, dehiscent monocar ps, and seeds with bilobed arils, A cladogram provides an explicit hyp othesis of intergeneric relationships in the tribe. The new combinatio n Onychopetalum periquino, based on Trigynaea periquino, is made.