S. Adalsteinsson et al., BROWN COAT COLOR IN ICELANDIC CATTLE PRODUCED BY THE LOCI EXTENSION AND AGOUTI, The Journal of heredity, 86(5), 1995, pp. 395-398
Inheritance of the colors black, brown, and red in Icelandic cattle wa
s studied. The three colors are produced by two loci, Extension (E) an
d Agouti (A), with three alleles at the E locus: E(d) for dominant bla
ck, E(+), intermediate, which allows expression of A locus alleles; an
d e for recessive red color. Two alleles are postulated at the A locus
: A(+), producing brown, and a, producing recessive black (nonagouti)
when homozygous in E(+)/- animals. The dominant and recessive types of
black are indistinguishable from each other phenotypically. The A all
eles are only able to express their effect in E(+)/- genotypes. The E
and A loci in cattle are postulated to be homologous to the E and A lo
ci in the mouse.