The hominoid Primates were first recogni sed in the early Miocene laye
rs of the African rift-valley in Eastern Africa. The genus Proconsul i
s a goad ancestral morphotype for all the superfamily. Proconsul was a
generalized arboreal ape without any specialisations of the recent ap
es. It is the starting point of an important radiation into the Early
and Middle Miocene of Africa. During the Middle Miocene the African co
ntinent came into contact with the Eurasian plate through the Arabian
plate and a land way was opened between the two continents and some fa
unal migrations occured at that time. Some lineages of hominoid Primat
es went to Europe and Asia. One of them, Sivapithecus, was found in As
ia, particularly in late Miocene deposits of the Siwalik hills; it sha
res derived characters with the recent Pongo and, in spite of some pec
uliar features, can be placed in the same clade which is the Pongidae
family. Another one was found in Northern Greece and it belongs to the
genus Ouranopithecus. The study of several teeth and jaws showed that
it shares derived characters with the australopithecines and must be
classified with the hominids. The recent discovery of a large part of
a face corroborates that Ouranopithecus is very far from Sivapithecus.
A supra-orbital torus is present but not projecting like those of the
African apes; the orbits is large; the nasal aperture is broad and th
e sides converge slightly toward the nasal bones which an flattened an
d narrowed at their proximal side. The skull of Ouranopithecus differs
from those of the other Miocene apes (Sivapithecus, Lufengpithecus or
Dryopithecus) and it fits better with Australopithecus afarensis. We
consider the Macedonian species as the fore-runner of the Plio-pleisto
cene hominids.