Growth in animals is defined as accretion of protein, fat, and bone. A
lthough growth typically is measured as the change in Live weight, nut
rient retention is estimated more precisely by measuring empty body we
ight and composition, whereas production economics are measured ideall
y through carcass weights and quality. As a percentage of Live weight
gain, carcass weight gain usually is a much higher percentage during t
he feedlot phase than during the growing phase of production because d
ressing percentage (ratio of carcass:live weight) increases with matur
ation and is greater with concentrate than with roughage diets. At a g
iven fraction of mature body size (maximum body protein mass), body fa
t percentage seems to be a constant. Mature size may be altered geneti
cally and nutritionally. Protein accretion declines to zero when cattl
e reach their mature body size (approximately 36% fat in empty body we
ight in modern cattle) even though mature animals can continue to accr
ete fat. Although fat accretion can be reduced by limiting the supply
of net energy, rate of fat accretion by finishing steers given ad libi
tum access to high-concentrate diets seems to reach a plateau at appro
ximately 550 g daily. Protein mass, in contrast, increases in proporti
on to empty body weight. The protein:fat ratio of the carcass can be i
ncreased through increasing mature size, by administering hormones or
hormonal modifiers, by limiting energy intake during the growing perio
d or finishing period, or by slaughtering cattle at an earlier stage o
f maturity. Energetically, efficiency of accretion of fat is approxima
tely 1.7 times that of protein. But because more water is stored with
deposited protein than with deposited fat, lean tissue gain is four ti
mes as efficient as accretion of fat tissue. Conversion of protein to
fat is very inefficient, suggesting that excess protein is utilized in
efficiently.