Fa. Coyle, A REVISION OF THE FUNNELWEB MYGALOMORPH SPIDER SUBFAMILY ISCHNOTHELINAE (ARANEAE, DIPLURIDAE), Bulletin of the American Museum of Natural History, (226), 1995, pp. 3-133
The tropical diplurid spider subfamily Ischnothelinae is revised for t
he first time. The monophyly of this subfamily is supported by six put
ative synapomorphies: an elongate cymbial apophysis, two rows of cheli
ceral teeth, a procurved transverse fovea, collariform trichobothrial
bases, fused silk spigots, and long, tapering, pseudosegmented termina
l posterior lateral spinneret articles. Five genera are recognized: Is
chnothele Ausserer (with 11 species in South America, Central America,
and the Antilles), Andethele, new genus (with 3 species in the Andes
Mountains of Peru), Lathrothele Benoit (with 4 species in Africa), Ind
othele, new genus (with 4 species in India and Sri Lanka), and Thelech
oris Karsch (with 2 species in Africa and Madagascar). Phylogenies (ba
sed on cladistic analyses using two different sets of outgroups), biog
eographic hypotheses, a key, diagnoses, descriptions, tables of quanti
tative character values, illustrations, analyses of variation, natural
history information, locality records, and distribution maps are pres
ented for the 24 recognized species. The cladistic analyses provide st
rong support for the monophyly of the Old World branch of the subfamil
y, the genera Andethele and Thelechoris, and the Greater Antillean and
South American (goloboffi) clades of Ischnothele, and weaker or ambig
uous support for the monophyly of Ischnothele, Lathrothele, and Indoth
ele. Ten species are newly described: Ischnothele jeremie, I. garcia,
I. huambisa, and I. goloboffi; Andethele lucma, A. huanca, and A. tarm
a; Indothele lanka, I. rothi, and I. mala. Fourteen specific names are
newly synonymized: Ischnothele ecuadorensis Schmidt with Ischnothele
digitata (O. P.-Cambridge); Entomothele pusilla Simon, Thelechoris zeb
rina Simon, Thelechoris funesta Fischel, Thelechoris obtusa Fischel, a
nd Ischnothele sexpunctata Bucherl, da Costa, and Lucas with Ischnothe
le caudata Ausserer; Ischnothele siemensi F. O. P.-Cambridge with Isch
nothele guianensis (Walckenaer); Ischnothele indigens Vellard, Ischnot
hele zorodes Mello-Leitao, Ischnothele campestris Schiapelli and Gersc
hman, Ischnothele affinis Schiapelli and Gerschman, and ischnothele cr
anwelli Gerschman and Schiapelli with Ischnathele annulata Tullgren; L
athrothele marmoratus Benoit with Lathrothele grabensis Benoit; and Th
elechoris karschi Bosenberg and Lent with Thelechoris striatipes (Simo
n). Several species previously placed in ischnotheline genera are not
ischnothelines: Ischnothele annectens (Bertkau), Ischnothele simplicat
a Saito, Ischnothele strandi Spassky, Ischnothele lineata (Karsch), Is
chnothele indicola Tikader, and Thelechoris australis Purcell. Evidenc
e is presented that suggests that Ischnothele longicauda and Lathrothe
le grabensis may each consist of more than one species and that Thelec
horis rutenbergi and T. striatipes may be conspecific.