Living systems theory (LST) generates a large set of hypotheses that a
re thought to be applicable to the following eight levels of living sy
stems: cell, organ, organism, group, organization, community, society,
and supranational system. Many of the disciplines from which LST has
derived its cross-level propositions are concerned not only with the p
roperties of individual systems, but also with interactions between th
em. This paper focuses on linked systems and the linkages between them
rather than on individual systems. Intersystem relationships at vario
us levels of living systems are examined for cross-level commonalities
. The paper pays particular attention to dyads as the building blocks
of quasi-living systems and higher-level living systems. Shared templa
te elements, a common destiny, joint activity, and exchange of resourc
es between and among systems are found to be common, cross-level facto
rs in the bonding of systems.