Type and additional specimens of species of Oryctocephalus, Lancastria
and Oryctocara are described. Oryctocephalidae have a rostral-hyposto
mal plate, and hence belong in the Order Corynexochida. Characteristic
of the family is the development of lateral glabellar furrows as pits
situated inside the axial furrow, and the absence of a border on the
pygidium. Two subfamilies are used: Oryctocephalinae, having genal and
pleural spines; Oryctocarinae lacking such spines. Species of both gr
oups are found in late Lower to mid Middle Cambrian rocks of North Ame
rica, Siberia, Asia and Australia, and rare examples occur in South Am
erica and Antarctica. Oryctocephalids are typical of deposits in outer
shelf and slope areas, in which completely or partially articulated e
xoskeletons are characteristic. Arrangements of sclerites in such spec
imens of Oryctocephalus burgessensis are considered to result from bur
ial events, not from manoeuvres during exuviation. Compaction and tect
onic distortion of specimens in these fine-grained elastic rocks rende
rs questionable some of the characters used in taxonomy.