Forested landscapes often show very well-pronounced heterogeneity in t
he factors that control evapotranspiration, runoff production and carb
on assimilation at a variety of length scales. In hilly or mountainous
environments, strong contrasts in net radiation, available soil water
, soil structure and stand characteristics can produce a large varianc
e in both the meteorological drivers and surface resistance to carbon
and water exchange with the atmosphere over distances measured in tens
of metres. Because of the strong nonlinearities characterizing the in
fluence of the environmental variables on surface resistance (particul
arly available soil water), the parametrization of surface process mod
els with mean values of the environmental variables and no distributio
n often leads to significant bias in areal average carbon and water fl
ux. However, it is often not feasible to incorporate directly the full
distribution and patterns of the landscape for regional-scale models.
Continental- and subcontinental-scale vegetation data sets currently
being collected by synoptic-level satellites (e.g. the Advanced Very H
igh Resolution Radiometer, AVHRR) do not capture the large proportion
of landscape variability that exists below the resolution of the senso
rs. This paper explores the impacts of various landscape representatio
n schemes that retain a range of detail in the description of land sur
face form and processes on simulated areal average evapotranspiration,
runoff production and net carbon exchange with the atmosphere. Specif
ic comparison is made of schemes that attempt to incorporate the topog
raphic structure, soil and vegetation distributions of a region with s
chemes that sample the surface at levels similar to current coarse-res
olution satellites. For strongly heterogeneous basins (mountainous top
ography), it is found that spatial variations in available soil water
can have significant effects on areal averaged carbon and water flux r
ates, particularly under drying conditions, whereas the spatial variat
ions in radiation, temperature and humidity over the terrain appear to
have a lesser impact.