This paper presents an historical review of the mechanisms of pollen c
ollecting by the Gymnosperm ovules. Nucellar and/or epidermal secretio
ns are involved in these mechanisms. Recently, the morphology, histolo
gy and cytology of the secreting tissues in ovules and nucelli have be
en explicited for some species using electron microscopy (transmission
and scanning E.M.). The pollination drops were analysed for many spec
ies. However, the pollen-ovule relations remain much less known than t
he pistil-pollen relations in Angiosperms.