Recent progress in the study of short-distance (cell-to-cell) movement
of plant virus, facilitated by 'movement proteins', has led to a resu
rgence of interest in long-distance virus transport in the phloem. Rel
atively little is known about phloem-specific barriers to virus moveme
nt or about the form in which virus enters, travels within and exits t
his tissue. Progress in understanding virus and photoassimilate transp
ort is limited by a paucity of information on the substructure and pro
perties of plasmodesmata at specific interfaces. The direction of viru
s movement, once it has entered the phloem, can be understood by follo
wing photoassimilate translocation, a complex and dynamic process infl
uenced by plant growth, development and vascular topology.