LARVAE OF TRICHOPHYA AND PHYLOGENY OF THE TACHYPORINE GROUP OF SUBFAMILIES (COLEOPTERA, STAPHYLINIDAE) WITH A REVIEW, NEW SPECIES AND CHARACTERIZATION OF THE TRICHOPHYINAE

Larvae of the staphylinid subfamily Trichophyinae are described for th e first time based on larvae of a new species of Trichophya from the s outhwestern United States. Adults and larvae of the new species, Trich ophya texana Ashe and Newton (type locality Texas, Brewster Co., Big B end National Park), are described and illustrations of both provided. Also given are a key for separation of the Nearctic species of Trichop hya, a checklist of the known World fauna of the Trichophyinae (includ ing first report of the genus from Mexico and Guatemala), and a charac terization of the subfamily Trichophyinae based on both larvae and adu lts. The relationships of major genera and higher taxa in the tachypor ine group of staphylinid subfamilies are analysed cladistically using larval characters. No larval characters were found that provide eviden ce for the monophyly of the tachyporine group; no evidence was found f or the monophyly of the Tachyporinae; Charhyphus, Olisthaerus and Phlo eocharis (Phloeocharinae + Olisthaerinae) form a monophyletic group; t he Trichophyinae and Habrocerinae are sister groups and together proba bly are the sister group to the Aleocharinae; the Aleocharinae are con firmed to be monophyletic based on larval characters; and Gymnusa + De inopsis form the sister group to the remainder of the Aleocharinae.