The shrub Monotoca is widespread in eucalypt forests. Monotoca-type po
llen is distinctive with a tetrad of one large functional cell and thr
ee small aborted cells. Other variations on the tetrad, with equal siz
ed cells, the normal pollen type for the family, cannot be confused wi
th Monotoca. Fossil Monotoca pollen first appears in the Mid Miocene,
when the climate was becoming drier, rainforest was contracting and sc
lerophyll forests were expanding.