Rw. Spear et al., LATE QUATERNARY HISTORY OF LOW-ELEVATION AND MID-ELEVATION VEGETATIONIN THE WHITE MOUNTAINS OF NEW-HAMPSHIRE, Ecological monographs, 64(1), 1994, pp. 85-109
Pollen and plant macrofossil analyses of sediments from an altitudinal
series of lakes in the White Mountains of New Hampshire, USA were use
d to reconstruct the history of vegetation on the mountain slopes and
to identify the factors responsible for vegetation change. Six sites,
Mirror Lake (213 m), Lost Pond (625 m), Little East Pond (793 m), Lone
some Lake (831 m), Carter Notch Pond (1004 m), and Lake of the Clouds
(1538 m), provide paleoecological records of changes in the altitudina
l limits of species, the species abundances within communities, and th
e vegetation zones. These are supplemented by previously published dat
a from three high-elevation sites (Spear 1989). Although past plant co
mmunities were different from modern ones, differentiation of vegetati
on along the slopes has always existed. At low elevations the sequence
of vegetation change was: 13 700-11 500 yr BP, tundra; 11 500-9000 yr
BP, transitional mixed-conifer woodlands of first spruce (Picea) and
then fir (Abies balsamea), larch (Larix laricina), poplar (Populus), a
nd paper birch (Betula papyrifera); 9000-7000 yr BP, forests dominated
by pine (Pinus) and oak (Quercus); 7000 yr BP-present, mixed-hardwood
forests. No late-glacial paleoecological records exist at mid-elevati
on sites (700-1200 m). The steep slopes at these elevations stabilized
by 10 000 yr BP and an early woodland of spruce was replaced by fir,
larch, and paper birch. The altitudinal limits of both white pine (Pin
us strobus) and hemlock (Tsuga canadensis) expanded to mid-elevations
during the mid-Holecene, suggesting great warmth 6000-4000 yr BP. The
modern spruce/fir forests of mid-elevations became established 2000 yr
ago as spruce expanded at all elevations, suggesting cooler, moister
climate similar to today. A tree line dominated by balsam fir and blac
k spruce (Picea mariana) was established at its modern position 10 000
yr ago and has varied little since then, although it appears to have
been slightly higher than now during the early Holocene. Changes in th
e vegetation at low and mid-elevations have not been synchronous with
those at tree line and result from a more complex set of environmental
factors and climatic variables acting on several different species. A
t low and mid-elevations disturbance by wind and frost action was impo
rtant during the late-glacial. Disturbance by fire was important durin
g the period of spruce woodland and later in the early Holocene pine a
nd oak forests. From 7000 yr BP to the present the primary factor dist
urbing New Hampshire forest was again probably wind, especially from 7
000 to 4000 yr BP when higher temperatures than present may have been
associated with increased frequency and intensity of thunderstorms and
tropical hurricanes. The vegetation history reveals that different la
pse rates have occurred along the mountain slopes. This provides evide
nce that the source of air masses reaching the White Mountains has var
ied. The boundary between alpine tundra and subalpine fir forest (tree
line) most likely has always been governed by temperature (summer ins
olation) and wind. The montane plant communities result from individua
l species response rather than community response to the numerous clim
atic forces that have affected the mountains over the past 14 000 yr.