Shear-wave splitting of S and SKS phases reveals the anisotropy and st
rain field of the mantle beneath the subducting Nazca plate, Cocos pla
te, and the Caribbean region. These observations can be used to test m
odels of mantle flow. Two-dimensional entrained mantle flow beneath th
e subducting Nazca slab is not consistent with the data. Rather, there
is evidence for horizontal trench-parallel flow in the mantle beneath
the Nazca plate along much of the Andean subduction zone. Trench-para
llel flow is attributable to retrograde motion of the slab, the decoup
ling of the slab and underlying mantle, and a partial barrier to flow
at depth, resulting in lateral mantle flow beneath the slab. Such flow
facilitates the transfer of material from the shrinking mantle reserv
oir beneath the Pacific basin to the growing mantle reservoir beneath
the Atlantic basin. Trench-parallel flow may explain the eastward moti
ons of the Caribbean and Scotia sea plates, the anomalously shallow ba
thymetry of the eastern Nazca plate, the long-wavelength geoid high ov
er western South America, and it may contribute to the high elevation
and intense deformation of the central Andes.