USAMBAROMYIA NIGRALA GEN-N, SP-N, AND USAMBAROMYIINAE, A NEW SUBFAMILY AMONG THE CHIRONOMIDAE (DIPTERA)

The male and female imagines of Usambaromyia nigrala gen. n., sp. n. a re described from the West Usambara Mountains in Tanzania. The species differs from other known chironomids by having nearly completely blac k wings in both sexes, tibial spurs with lateral denticles making them Tanypodinae - like, dense cluster of scalpellate sensilla chaetica on anepisternum I, male gonostylus without megaseta, laterosternite fuse d with tergite IX forming a single annulus and female genitalia appare ntly with only one seminar capsule and spermathecal duct, well develop ed gonocoxite IX and setae on segment X. A new subfamily, the Usambaro myiinae is erected based on U. nigrala and conceived as forming the pl esiomorphic sister group of the Tanypodinae, the Podonomiinae and the Aphroteniinae combined. A subfamily diagnosis is given.