Extant ungulates can be divided into three dietary categories: browsin
g feeders, grazing feeders, and mixed feeders. Dietary adaptations can
be differentiated in extinct ruminants based upon tooth microwear ana
lysis as well as evaluation of premaxillary morphology. Tooth microwea
r shows that the extinct bovid Kipsigicerus labidotus from the Miocene
of Fort Ternan in Kenya (14 million years old) was most likely a graz
ing feeder, with mixed-feeder tendencies, while morphologically the pr
emaxilla most closely resembles that of a mixed feeder. Because the pa
leoenvironment at Fort Ternan was likely to have been forested, as sho
wn by paleosol isotopic studies, grazing in this particular ruminant e
volved within a forested environment preceding the origin of savanna.