M. Kottelat et Kkp. Lim, A REVIEW OF THE EEL-LOACHES OF THE GENUS PANGIO (TELEOSTEI, COBITIDAE) FROM THE MALAY PENINSULA, WITH DESCRIPTIONS OF 6 NEW SPECIES, The Raffles bulletin of zoology, 41(2), 1993, pp. 203-249
Ten species of Pangio occur in the Malay Peninsula: P. anguillaris, P.
cuneovirgata, P. doriae, P. kuhlii, P. malayana, P. oblonga, P. shelf
ordii, and the new species P. alcoides, P. filinaris, and P. piperata.
Pangio alcoides is distinguished by a high vertebral count, a very sm
all eye, and a peculiar colour pattern (usually white belly and black
dorsum); P. filinaris by a plain brown body and a nasal barbel, P. pip
erata (which also occurs in Sumatra) by a brown body with fine black d
ots forming small blotches along dorsal midline and a relatively low v
ertebral count. Three additional new species are described from Borneo
: P. alternans from the Mahakam basin, Borneo, is distinguished by its
colour pattern of alternating blotches on lower half of body and bars
and by its low vertebral count; P. incognito (which occurs syntopical
ly with P. shelfordii in Sarawak) by a low vertebral count and details
of coloration; P. pulla from Kalimantan Tengah, Borneo, by the absenc
e of pelvic fins, its colour pattern consisting of broad black bars an
d narrow interspaces, and a high vertebral count. Pangio malayana and
P. myersi (from southeastern Thailand) are removed from the synonymy o
f P. kuhlii and considered valid species; P. muraeniformis is consider
ed a synonym of P. shelfordii; P. borneensis is possibly a senior syno
nym of P. shelfordii but its status cannot be solved without new colle
ctions. Pangio shelfordii, P. cuneovirgata, P. doriae and P. superba a
re recorded for the first time from Sumatra. Neotypes are designated f
or Acanthophthalmus fasciatus Bleeker, 1860, Cobitis oblonga Valencien
nes, 1846, and A. javanicus Bleeker, 1860.