Notable among the biological changes of the middle part of the Maastri
chtian Age was a major pulse of extinction among inoceramid bivalves.
Analysis of the distribution of inoceramid shell fragments suggests th
at the population of burrowing organisms increased across the same str
atigraphic interval that the bivalve population decreased. A global re
organization of ocean circulation that resulted in cooler, better oxyg
enated bottom waters is proposed as an explanation for the observed ch
anges in the deep sea. Other observations regarding the mid-Maastricht
ian are consistent with this hypothesis.