Following principles used by A. J. Bateman, we identify the relationsh
ip between fecundity and mating success as the central feature in the
operation of mating systems. Using selection theory from the field of
quantitative genetics, we define the sexual selection gradient as the
average slope of the relationship between fecundity and mating success
and show how it can be estimated from data. We argue that sexual sele
ction gradients are the key to understanding how the intensity of sexu
al selection is affected by mate provisioning, parental investment, an
d sex ratio.