MAKING CONNECTIONS IN FOOD WEBS

Patterns in food web structure have provided an important, though cont entious, testing ground for ideas about the population dynamics and en ergetic of multispecies systems. One of the most debated of these patt erns is the apparent decrease in food web connectance as the number of species in a web increases. Several contrasting mechanisms that might determine food web connectance have been suggested. These mechanisms, in combination with new, food web data, suggest that the conventional pattern, and explanations for it, may be open to dispute. The true na ture of the relationship between connectance and species number has im plications for the explanations of other web patterns and for theories of food web structure, but a general explantation remains elusive.