B. Roth et Kc. Emberton, EXTRALIMITAL LAND MOLLUSKS (GASTROPODA) FROM THE DEEP RIVER FORMATION, MONTANA - EVIDENCE FOR MESIC MEDIAL TERTIARY CLIMATE, Proceedings of the Academy of Natural Sciences of Philadelphia, 145, 1994, pp. 93-106
Based on sedimentation patterns, lithology, and fossil vertebrates, th
e period from late Eocene to late early Miocene has been considered a
time of dry climate in the northern Rocky Mountains. Analysis of fossi
l land snails from strata of Arikareean (late Oligocene to early Mioce
ne) age from the Deep River Formation, Meagher County, Montana, howeve
r, indicates a mesic climate with at least 75 cm/yr precipitation and
suggests the presence of woodland with climatic requirements like thos
e of present-day mixed mesophytic forest. Several species belong to ex
tant genera whose modern range limits have been displaced eastward, of
f the North American cordillera. Among these, two new species of land
gastropod, Hendersonia stillmani (Helicinidae) and Euchemotrema (forme
rly Stenotrema) occidaneum (Polygyridae), are described.