Figs and fig wasps form one of the best known examples of species-spec
ific mutualism and coevolution. Recent experiments and observations ha
ve led to a better understanding of the evolutionary processes involve
d in the origin and maintenance of species interactions. The observed
fine-tuned traits involve not only coevolution but also selection acti
ng on only one of the partners. Furthermore, some of the 'fine-tuned t
raits' appear to be preadaptations - traits that existed before the mu
tualism was established.