Hl. Simpson et Mp. Denne, VARIATION OF RING WIDTH AND SPECIFIC-GRAVITY WITHIN TREES FROM UNTHINNED SITKA SPRUCE SPACING TRIAL IN CLOCAENOG, NORTH-WALES, Forestry, 70(1), 1997, pp. 31-45
The aim of this work was to quantify patterns of change in ring width
and specific gravity within trees of Picea sitchensis with ring number
across juvenile and mature wood, in relation to height in tree, origi
nal spacing, and crown dimensions. Five trees were sampled from each o
f three plots on a 52-year-old unthinned spacing trial at Clocaenog, N
orth Wales. The ring width of wood produced in early decades of the pl
antation was most strongly correlated with original spacing, while tha
t produced in later decades was more strongly correlated with branch d
iameters of the upper crown. In later decades, trees originally at the
widest spacing had higher specific gravity than those originally at c
loser spacing, presumably associated with self-thinning of the narrowe
r-spaced unthinned plots. Differences in amount of juvenile wood betwe
en spacing plots were less marked than those reported from the same pl
ots when harvested 7 years earlier; it is suggested that as trees surv
iving after self-thinning are likely to be those with a competitive ad
vantage from an early stage, differences between plots in amounts of j
uvenile wood may become less apparent in trees harvested later in the
rotation than in those sampled earlier. Underlying patterns of variati
on in ring width and specific gravity across the tree were found to be
modified by a progressive drift with height in the tree. Linear regre
ssions between specific gravity and ring width also varied in a system
atic way; the intercept and slope of these regressions tended to incre
ase with ring number from the pith, and with height in tree at a speci
fied ring number. Equations are given as a basis for quantifying these
trends, but more data are needed from other sites to determine the ex
tent to which these equations represent trends inherent to cambial and
apical ageing, as distinct from influence of changing environment aro
und the trees.