A mean lift coefficient quasi-steady analysis has been applied to the
free flight of the dragonfly Sympetrum sanguineum and the damselfly Ca
lopteryx splendens. The analysis accommodated the Saw and acceleration
s involved in free flight, For any given velocity or resultant aerodyn
amic force (thrust), the damselffy mean lift coefficient was higher th
an that for the dragonfly because of its clap and fling, For both spec
ies, the maximum mean lift coefficient <(C)over bar (L)> was higher th
an the steady C-L,C-max Both species aligned their strokes planes to b
e nearly normal to the thrust, a strategy that reduces the <(C)over ba
r (L)> required for flight and which is different from the previously
published hovering and slow dragonfly flights with stroke planes steep
ly inclined to the horizontal, Owing to the relatively low costs of ac
celerating the wing, the aerodynamic power required for flight represe
nts the mechanical power output from the muscles, The maximum muscle m
ass-specific power was estimated at 156 and 166 W kg(-1) for S, sangui
neum and C, splendens, respectively, Measurements of heat production i
mmediately after flight resulted in mechanical efficiency estimates of
13% and 9% for S, sanguineum and C, splendens muscles, respectively.