J. Cluttonbrock et al., CHANGES IN THE SKULL MORPHOLOGY OF THE ARCTIC WOLF, CANIS-LUPUS-ARCTOS, DURING THE 20TH-CENTURY, Journal of zoology, 233, 1994, pp. 19-36
Measurements of a large series of skulls of the Arctic wolf Canis lupu
s arctos, have shown that since 1930 there has been an overall reducti
on in the size of the skulls, together with widening of the cranium, s
hortening of the facial region, and reduction in size of the teeth. Th
is suggests that hybridization and subsequent introgression occurred w
ith huskies (Canis familiaris) during the 1930s, which is consistent w
ith historical accounts. Since 1950 there has been a reversion in skul
l morphology to a more 'wolf-like' form, suggesting that hybridization
is no longer occurring. The skull of a wolf/dog hybrid is intermediat
e in size between the skulls of wolves and huskies but its shape is al
lometrically dissimilar. Skulls of wolves from the period 1930-50 are
more similar to the skull of this hybrid than in the other time period
s. The skull of a male canid from a carcass collected on Ellesmere Isl
and and presented to the Natural History Museum, London, in 1986 was a
t first thought to be from a wolf/dog hybrid but analyses of the measu
rements show that it is more likely to be from an Arctic wolf with sev
ere abnormalities to the jaws.