A. Sikder et Ab. Roy, MULTIPLE CYCLIC SETS CONNECTING SADDLE-POINTS AND LIMIT-CYCLES OF A 4-SPECIES SYSTEM AND ITS UNIFORM PERSISTENCE, Mathematical biosciences, 121(2), 1994, pp. 141-154
Considering a model of a generalized Gause-type four-species system of
two predator-prey pairs linked by competition, the authors derive suf
ficient conditions for the existence of three different invariant cycl
ic sets connecting saddle points and limit cycle(s) on the boundary of
its state space. Also discussed is the uniform persistence of the mod
el in the presence of such cyclic sets.