S. Lovari et M. Appollonio, ON THE RUTTING BEHAVIOR OF THE HIMALAYAN GORAL NEMORHAEDUS-GORAL (HARDWICKE, 1825), Journal of ethology, 12(1), 1994, pp. 25-34
The rutting behaviour of the Himalayan goral Nemorhaedus goral was stu
died in an area of the Himalayan foothills, Himachal Pradesh, Northern
India, from October 13 to 25, 1989. Twenty seven qualitatively differ
ent behaviour patterns were observed. Males showed a more extensive re
pertoire (25 patterns, 14 performed only by this sex) than did females
(12 patterns, 1 of them-Reactive urination-unique). The Head butt and
the Intentional head butt were the most frequently shown behaviour pa
tterns of both sexes, and Naso-nasal contact followed. Males interacte
d mainly with females and usually seemed to ignore each other. Courtsh
ip activities peaked on October 22-25, when mounts were also recorded.
''Following'' was the only mating system observed. Pairs sometimes co
urted at short distances from each other, without eliciting any appare
nt aggressive reaction. No evidence of territoriality was found throug
hout our study period. The goral behaviour appears to be more closely
related to that of Oreamnos (in quality and quantity of behaviour patt
erns, as well as the mating system) than to that of Capricornis.