This article examines the mechanisms of subsistence adaptation of two
Papua New Guinea populations, the Metroxylon sago-depending lowland Gi
dra and the taro-monoculture Mountain Ok, surviving in low population
densities of 0.5 and 1.4 persons per km2. Observation of the groups' l
and use systems strongly suggests that their population densities have
not been far below the carrying capacity, although the territory of e
ach population is markedly heterogeneous. Both groups have maintained
their sustainable food production not only for resource management but
also for survival at a population level, either expanding their terri
tory or changing the sustainable level in tandem with changes of subsi
stence system.