S. Jose et al., STRUCTURAL, FLORISTIC AND EDAPHIC ATTRIBUTES OF THE GRASSLAND SHOLA FORESTS OF ERAVIKULAM IN PENINSULAR INDIA, Forest ecology and management, 65(2-3), 1994, pp. 279-291
The high elevation shola-grassland vegetation types of the Western Gha
ts apparently remained in a stable equilibrium for many decades, imply
ing that both vegetational types are highly developed and attained sta
bility under the same climatic regime. Several authors considered thes
e two distinct vegetation formations occuring in juxtaposition, as cli
max formations. According to the Clementsian view of climatic climax,
however, two distinctly different vegetation types cannot form climati
c climaxes under the same regional climate. Possibly some edaphic or b
iotic factors are responsible for rendering stability to the grassland
s in this system. We tested the following three hypotheses in this con
text. ( 1 ) The physico-chemical properties of the soil such as depth,
organic matter, nutrient status and water holding capacity are more f
avourable in the depressions than in the exposed surfaces, which in tu
rn might be responsible for the development of the woody vegetation in
the depressions. (2) The shola forests exhibit similar structural att
ributes and a higher floristic diversity compared with the medium and
low elevation wet evergreen forests of the Western Ghats region. (3) T
he shola trees are characterised by a low regeneration capacity, thus
probably making it a shrinking resource base. With its characteristic
deep fertile soil and high moisture holding capacity, the shola forest
s may remain in the same steady climax state, provided anthropogenic a
nd other catastrophic disturbances do not destroy them. The grasslands
were, however, characterised by generally shallow soils, low water ho
lding capacity and low site nutrient capital. Hence, the assumption th
at grasslands are steady state vegetations maintained by edaphic facto
rs, holds good. Structural and floristic elements of the shola forest
revealed a very high floristic richness and diversity, probably the hi
ghest in the Western Ghats region. A total of 942 stems of at least 10
cm girth at breast height (GBH) and belonging to 5 3 species were enc
ountered in the 5000 m2 sampling area (basal area 48 M2 ha-1). Contrar
y to the widely held dogma that shola forest does not possess adequate
regeneration potential, the present study reveals that under the fore
st cover, profuse regeneration of almost all of the overstorey species
occur. However, regeneration characteristics were different along the
margins and in open grasslands, possibly as a result of differences i
n ecological conditions.