An early tetrapod fossil from the Upper Devonian of Pennsylvania (Cats
kill Formation) extends the temporal range of tetrapods in North Ameri
ca and suggests that they attained a virtually global equatorial distr
ibution by the end of the Devonian. Derived features of the shoulder g
irdle indicate that appendicular mechanisms of support and propulsion
were well developed even in the earliest phases of tetrapod history. T
he specialized morphology of the pectoral skeleton implies that the di
versity of early tetrapods was great and is suggestive of innovative l
ocomotor patterns in the first tetrapods.