Sjr. Woodward et Gc. Wake, A DIFFERENTIAL-DELAY MODEL OF PASTURE ACCUMULATION AND LOSS IN CONTROLLED GRAZING SYSTEMS, Mathematical biosciences, 121(1), 1994, pp. 37-60
A grazing population dynamics model is proposed where organisms in a g
razed population have a fixed life span. The motivating context is tha
t of ruminants grazing grass-dominant pasture. The model takes the for
m of a differential-delay equation in which the rate of loss of pastur
e due to senescence at some time depends on the rate at which leaves a
re reaching maturity at that time. Comparisons are made with data from
a continuous grazing experiment due to Bircham and Hodgson (Grass and
Forage Science, 38:323-331, 1985), leading to a prediction of 21.9 da
ys for herbage life span. Predictions of herbage utilization are consi
stent with measured data. The model predicts lower senescence in sward
s in regrowth than in grazed swards at the same herbage mass. Solution
s and equilibria are obtained for the linear form of the model with co
ntinuous grazing pressure. Solutions and bounds are obtained for the l
inear model with intermittent grazing pressure, and its usefulness in
modeling grazed pastures is discussed. A delay model is a simple but p
owerful means of including the concept of fixed herbage life span in g
razing modeling. Questions of herbage life span and percentage utiliza
tion are naturally contained in the mechanism of a differential-delay
model. These are not so well handled by models that treat senescence o
f herbage empirically.