Ch. Dietrich, SYSTEMATICS OF THE LEAFHOPPER GENUS DRAECULACEPHALA BALL (HOMOPTERA, CICADELLIDAE), Transactions of the American Entomological Society, 120(2), 1994, pp. 87-112
A checklist and synonymy for the 25 species of Draeculacephala is pres
ented and an illustrated key based on discrete morphological character
s is provided for species identification. Three new Neotropical specie
s are described and illustrated. Previously unrecognized morphological
characters useful for delimiting species and species groups include t
he number of paleate setae on hind tarsomere I and the distribution of
erect setae at the base of the male pygofer. Weighted and unweighted
parsimony analyses of morphological characters of the species of Draec
ulacephala and selected species of the related cicadelline genera Helo
chara Fitch, Syncharina Young, and Xyphon Hamilton support the status
of Draeculacephala as a monophyletic group. Placement of D. floridana
Ball, type species of Carneocephala Ball, in Draeculacephala is also s
upported. Draeculacephala apparently originated in Mexico or the south
western U.S. and comprises three major lineages that diversified in is
olation: 1) D. clypeata Osborn and related Central and South American
species; 2) D. crassicornis Van Duzee and related species inhabiting n
orth temperate North America; and 3) the remaining species, which are
concentrated in the southeastern U.S.