Cj. Devilliers et Ac. Brown, SAND MOISTURE AS A FACTOR DETERMINING DEPTH OF BURROWING IN THE ONISCID ISOPOD TYLOS-GRANULATUS KRAUSS, South African journal of zoology, 29(2), 1994, pp. 186-188
Tylos granulatus, a sandy-beach isopod, prefers an environmental moist
ure range exceeding 3,4% but less than 13%. The depths to which the an
imals burrow are, at least partly, determined by the moisture gradient
in the sand. They are, however, incapable of burrowing into totally d
ry sand. Animals after their position in the sand in response to chang
es in moisture content so as to ensure exposure to suitable conditions
.