THE EXTRAORDINARY LARVAL CHARACTERS OF CARENUM-BONELLI AND THEIR BEARING ON THE PHYLOGENY OF THE SCARITITAE (COLEOPTERA, CARABIDAE)

Larvae of the Australian fossorial carabid Carenum anthracinum Macleay are described from reared material and compared with the reared larva of C. elegans Macleay and eight different unassociated larval types b elonging to the complex. These larvae exhibit an extraordinary range o f character states indicating clearly that their placement in the trib e Scaritini cannot be retained. An examination of adult characters and , in particular, the fore tibial dentition confirms that Carenum and a llied genera form a separate tribe, the Carenini, which appears to be a sister group of the remaining Scaritini (including Clivinina).