A mechanism for phloem translocation is proposed based on a loading-un
loading hypothesis. The forces are many local tandem moving pressure g
radients. Evidence to support this hypothesis is developed from the pu
blished literature along with some newly presented data. This mechanis
m reconciles phenomena of pressure-flow, contractile protein, and elec
trical hormonal control in an hypothesis which is essentially active a
nd ultimately symplastic for both loading and unloading of sieve tubes
. It has permitted predictions to be made and tested successfully in s
ome twenty experiments to date.