A. Kratochwil et A. Schwabe, COINCIDENCES BETWEEN DIFFERENT LANDSCAPE ECOLOGICAL ZONES AND GROWTH FORMS OF CEMBRAN PINE (PINUS-CEMBRA L) IN SUB-ALPINE HABITATS OF THE CENTRAL ALPS, Landscape ecology, 9(3), 1994, pp. 175-190
The 'forgotten' depots of the European Nutcracker (Nucifraga c. caryoc
atactes) often 'lead' to the development of tufts of Pinus cembra. In
many cases the other individuals of such tufts are not suppressed by t
he fittest one, rather there is an intraspecific coexistence up to the
senescent stage of the trees. There are fusions of separate trunks, a
nd so frequently the individual history of older trees can only be rec
onstructed by studying sutures, crown structures or trunk cross sectio
ns. Different types of trunk fusions are worked out. By means of trans
ect counting the occurrence of these 'multiple trunk trees' is documen
ted quantitatively in different landscape ecological zones of the Enga
din region (the Grisons, Switzerland). The data base is 3024 counted m
icrosites of Pinus cembra individuals arising from seeds, including 52
72 living individuals. These 'multiple trunk trees' significantly play
an important role in the landscape ecological zones of recent glacier
recession and at the alpine timberline. Their growth forms have a hig
her biomechanical stability.