Strigiformes have a very long palaeontological history and were much m
ore diversified in the past than they are at present. Berruornis orbis
antiqui gen. et sp. nov., from the Upper Palaeocene (Thanetian) of the
Reims area, is a large owl, with a stout tarsometatarsus. It belongs
to the Sophiornithidae, previously described from the Upper Eocene to
Upper Oligocene of the Phosphorites du Quercy, France. The comparison
of the distal part of the tarsometatarsus in different extinct forms s
hows a general trend towards a progressively more semizygodactyl foot,
with an internal trochlea more posteriorly oriented, and an external
trochlea more medially incurved; correspondingly the size of the inter
nal trochlea compared with the middle trochlea decreases.