An Ordovician Promopalaeaster (Echinodermata: Asteroidea) wrapped abou
t a Cuneamya? (Mollusca: Pelecypoda) in the extraoral feeding posture
characteristic of Jurassic to modern members of the Asteriidae documen
ts an early origin for this behavioral complex. Modern asteriids are c
onvergent on Promopalaeaster, there is no direct phylogenetic linkage
between the two. This fossil occurrence, combined with the success of
modern Asteriidae, demonstrates that biological evolution and geologic
al change need not outmode complex life habits. The fossil supports th
e notion of asteroids as Paleozoic-type predators, and its existence s
uggests that asteroids were not significant contributors to changing f
aunal structures in shelf seas during the Phanerozoic.