Three 27-month-old infant gorillas living with their mothers and a sil
verbacked male were separated to a cage for 24 weeks. The infants init
ially showed threat responses and increased locomotion, characteristic
of the protest stage of anaclitic depression in children. Within seve
ral days, these were replaced by dorso-ventral contact among the infan
ts as well as self-holding and fetal positioning. Additionally, social
and solitary play and object examination occurred at lower levels thr
ough separation than in the pre-separation condition. These changes we
re characteristic of the despair stage of separation. There was a subs
tantial recovery of many infant nonsocial and social behaviors in the
later months of the separation. Upon reunion, the infants did not imme
diately engage in attachment behaviors with their mothers, and spent m
ore time in contact with each other than with their mothers for the fi
rst several days, indicating detachment. Following this, there was an
increase in mother-infant attachment behaviors. (C) 1994 John Wiley an
d Sons, Inc.