The giant deer, Megaloceros giganteus, is best known from its fossil o
ccurrences in Ireland around 11000 years ago, but has a history across
Europe and Western Asia spanning 300000-400000 years. This paper repo
rts a biometric study of variation and evolution in the giant deer thr
ough its history. Most early populations were as large in body size as
the Irish sample, but some were distinctly smaller and others had mar
kedly shorter limbs. Thickened skull and mandibular bones, especially
marked in males, are constant throughout the species' history. Some ea
rlier populations had antlers which were relatively smaller, more upri
ght and differing in other details from the celebrated Irish specimens
. Observed variation in body and antler form is discussed in terms of
changing palaeoenvironments. Interpretation of the fighting or display
roles of M. giganteus antlers needs to take account of their differin
g forms through time.