P. Tassy, GAPS, PARSIMONY, AND EARLY MIOCENE ELEPHANTOIDS (MAMMALIA), WITH A REEVALUATION OF GOMPHOTHERIUM ANNECTENS (MATSUMOTO, 1925), Zoological journal of the Linnean Society, 112(1-2), 1994, pp. 101-117
A new analysis of the morphology of the cranium of the early Miocene s
pecies Gomphotherium annectens (Matsumoto, 1925) alters significantly
previous hypotheses on the differentiation of Miocene Elephantoidea. T
he gomphorheres, excluding shovel-tusked mastodonts and choerolophodon
ts, are known to be a paraphyletic grouping: a stem group for elephant
ids+stegodontids. From parsimony analyses it seems very likely that ea
rly Miocene Old World so-called gomphotheres (Gomphatherium 'annectens
group') are not closely related to other gomphotheres.