Mitani and Rodman (1979) showed that a simple measure of the defendabi
lity of the range area could differentiate fairly successfully between
territorial and non-territorial species of primates. Their analysis h
as, however, been much criticised on the grounds that it considered on
ly the day journey length and the diameter of the range area. We devel
op a number of more realistic indices of territory defendability that
take into account both the length of the boundary to be defended and t
he detection distance as well as linear measures of range size. These
indices (a) discriminate between territorial and non-territorial speci
es more successfully than the Mitani-Rodman index and (b) allow the re
sults to be extended to species which do not forage in cohesive groups
. However, our results suggest that the Mitani-Rodman analysis was in
many ways correct.