THE PHYLOGENY AND CLASSIFICATION OF THE DISEASE (ORCHIDOIDEAE, ORCHIDACEAE)

The subtribal classification of the Disease (Orchidoideae) is reviewed in light of the available morphological, leaf anatomical, and palynol ogical data. These data are critically assessed, and the more prominen t features are illustrated. The data are analyzed cladistically, and t he robustness of the various components of the most parsimonious tree is assessed by a bootstrap analysis. Based on the cladistic analysis a nd the bootstrap analysis, a new classification is proposed for the Di seae. The results of the bootstrap analysis are used to establish the nodes at which formal taxa should be recognized. This classification r ecognizes five monophyletic subtribes: the Satyriinae, Disinae, Brownl eeinae (a new subtribe), Huttonaeinae, and Coryciinae. It is suggested that Brownleea, the only genus of the Brownleeinae, may be of hybrid origin, as it shares the autopomorphies of the Disinae and Coryciinae. Huttonaea is shown to be more closely related to the Diseae than to t he Orchideae, and is consequently included as a subtribe of the Diseae . The new classification is formally presented, and a key to the gener a is provided.