We measured nipple sizes of 29 captive wolves (Canis lupus), of known
breeding histories, throughout the year and tested distinctions among
various known breeding statuses of 20 wild wolves examined in northeas
tern Minnesota from May through September. For ca. 8 mo of the year on
ly breeders and nonbreeders can be classified. Distinctions between cu
rrent and former breeders were not reliable.