A single specimen of the early Middle Cambrian (Ordian) orthothecimorp
h hyolith Guduguwan hardmani with preserved gut filling has been disco
vered in the Montejinni Limestone, Wise Basin, northern Australia. Thi
s represents only the eighth such occurrence worldwide. The specimen i
s unique among orthothecimorphs in retaining the gut filling in combin
ation with the external mould of the operculum and invites revision of
the accepted soft-part reconstruction in this group, whereby the oper
cular lateral processes are now recognized to be ventral. Thus orthoth
ecimorph conchs, if horizontal in life, would have been oriented ventr
al side down, while hyolithomorphs adopted the reverse orientation: ho
rizontal, ventral side up.