Inferred phylogenetic relationships within the Brachiopoda have long b
een accepted as demonstrating repeated transitions from a phosphatic s
hell chemistry to a calcareous composition. This interpretation is ref
lected in the major subdivision of the phylum into the classes Inartic
ulata and Articulata. Cladistic analysis suggests, however, that the p
hosphatic-shelled and the calcareous-shelled Brachiopoda are sister gr
oups that have had consistently separated shell chemistries from early
in phylogenetic history. This separation is recognized in the class-r
ank divisions Lingulata and Calciata, the former of which includes the
new Subclass Lingulatea, whilst the latter includes the new Subclass
Craniformea together with the 'articulates' of previous classification
s.