H. Fery et An. Nilsson, A REVISION OF THE AGABUS-CHALCONATUS-GROUP AND ERICHSONI-GROUPS (COLEOPTERA, DYTISCIDAE), WITH A PROPOSED PHYLOGENY, Entomologica Scandinavica, 24(1), 1993, pp. 79-108
The Holarctic chalconatus- and erichsoni-groups, with 19 and three spe
cies respectively, are revised. The following seven new species are de
scribed: Agabus balkei from Siberia, Agabus jaechi from Turkey, Agabus
larsoni from Canada, Agabus lenkoranensis from Azerbaijan, Agabus ped
erzanii from Italy, Agabus samokovi from Bulgaria, and Agabus wewalkai
from Turkey. Lectotypes are designated for the following nominal spec
ies: Dytiscus chalconatus Panzer, 1796, Dytiscus nigroaeneus Marsham,
1802, Colymbetes montanus Stephens, 1828, Agabus neglectus Erichson, 1
837, Agabus subtilis Erichson, 1837, Agabus gagates Aube, 1838, Agabus
altaicus Gebler, 1848, Agabus erichsoni Gemminger & Harold, 1868, Gau
rodytes lutosus Crotch, 1873, Agabus fuscoaenescens Regimbart, 1877, A
gabus aenescens Poppius, 1905, Agabus satunini Zaitzev, 1913, and Agab
us melanocornis Zimmermann, 1915. The following new synonyms are estab
lished: Agabus altaicus Gebler, 1848 = Agabus subtilis Erichson, 1837;
Agabus melanocornis Zimmermann, 1915 = Colymbetes montanus Stephens,
1828; Gaurodytes bulgaricus Csiki, 1943 = Dytiscus conspersus Marsham,
1802; Agabus skiathos Hinterseher, 1981 = A gabus pseudoneglectus Fra
nciscolo, 1972; and Agabus maestroae Fresneda & Hernando, 1987 = Agabu
s albarracinensis Fery, 1986. Agabus aenescens Poppius, 1905, and A. s
atunini Zaitzev, 1913, are recognized as valid species (sp. rev.). The
chalconatus-group is divided into four subgroups and the phylogeny of
the group is reconstructed. The relationships of the chalconatus- and
erichsoni-groups with other Agabus species-groups and with Ilybius Er
ichson are discussed.